The drama explores the Philippine mythology surrounding aswangs, Juan dela Cruz (Coco Martin) is the son of mortal, Amelia (Mylene Dizon) and aswang (Albert Martinez). Juan is next in the lineage of tagapagbantays but unknown to him he bears aswang blood. The aswangs must not know of this fact for they will force him to join their clan, therefore he is brought up to do good deeds and believe in God to forget half of his very being. But the metal cross that is left by his mother before her death is stolen, the cross has the ability to shift into different weapons: a whip, a spear, an arrow, and a sword to fight against evil. In his quest to find his missing cross, he meets and falls in love with Rosario (Erich Gonzales) an undercover aswang-hunter.
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